Nouvel ichnogenre et nouvelle ichnoespèce de cubichnium, le Pygocephalichnium reidi, du site du patrimoine mondial de l’UNESCO des falaises fossilifères de Joggins, au Canada, et ichnotaxons connexes

Auteurs-es

  • Bailey C. Malay Laboratoires paléontologiques Steinhammer, Section de géologie/paléontologie, Département d’histoire naturelle, Musée du Nouveau-Brunswick, Saint John (Nouveau-Brunswick) E2K 1E5 CANADA
  • Matthew R. Stimson Laboratoires paléontologiques Steinhammer, Section de géologie/paléontologie, Département d’histoire naturelle, Musée du Nouveau-Brunswick, Saint John (Nouveau-Brunswick) E2K 1E5 CANADA
  • Olivia A. King Département des sciences de la terre, Université Carleton, Ottawa (Ontario) K1S 5B6 CANADA
  • Sal Faulkner Département des sciences de la mer, Centre des sciences de la mer de l’Université Memorial de Terre-Neuve, Logy Bay, (Terre-Neuve-et-Labrador) A1C 5S7 CANADA
  • Brian L. Hebert Fundy Treasures, 107, route 242, Joggins (Nouvelle-Écosse) B0L 1A0 CANADA
  • John H. Calder Département de géologie, Université Saint Mary’s, Halifax (Nouvelle-Écosse) B3H 3C3 CANADA
  • R. Andrew MacRae Département de géologie, Université Saint Mary’s, Halifax (Nouvelle-Écosse) B3H 3C3 CANADA
  • Spencer G. Lucas Musée d’histoire naturelle du Nouveau-Mexique, Albuquerque, Nouveau-Mexique 87104 ÉTATS-UNIS

DOI :

https://doi.org/10.4138/atlgeo.2024.009

Résumé

Le site du patrimoine mondial de l’UNESCO des falaises fossilifères de Joggins, en Nouvelle-Écosse, au Canada, est reconnu depuis longtemps pour sa paléobiodiversité considérable. L’enregistrement de l’ichnofossile à Joggins est moins connu que le fossile corporel. Les vastes collections ichnologiques du défunt citoyen scientifique Donald Reid comprennent un cubichnium évoquant la forme d’une crevette morphologiquement unique (trace au repos). L’ichnofossile est associé à une piste d’invertébré à peine visible menant à la trace au repos et permet son identification à titre de trace au repos d’invertébré. L’ichnofossile a été recouvré de la partie supérieure de la Formation de Joggins (à 876 m au-dessus de la base) et il a été découvert dans des grès ridés à grains fins, interprétés comme un vestige de la transition d’un assemblage de lithofaciès d’eau libre à un milieu mal drainé. La plaquette d’ichnofossile étudiée aux présentes comporte également des traces au repos d’invertébrés Selenichnites et Rusophycus préservés dans un hyporelief convexe; ces traces sont communément attribuées à des limules et des crustacés, respectivement. Elles se manifestent dans le même horizon stratigraphique que les pistes de Kouphichnium, interprétées comme des empreintes produites par des xiphosures. La tranche supérieure de la Formation de Joggins a auparavant livré des fossiles corporels de crevette Pygocephalus préservés dans des calcaires riches en matières organiques et des nodules sidéritiques-ferrugineux. Les fossiles corporels de Pygocephalus sont répandus à Joggins, mais aucun ichnofossile n’a été rattaché à cet invertébré, sans doute parce qu’aucun n’avait été identifié jusqu’à présent. L'ichnofossile nouvellement défini aux présentes à titre de Pygocephalichnium reidi est interprété comme une empreinte produite par une crevette Pygocephalus d’après ses similarités morphologiques avec des fossiles corporels connus de la Formation de Joggins et nous avançons que la morphologie de ce nouvel ichnofossile justifie la reconnaissance d’un nouvel ichnotaxon, le Pygocephalichnium reidi.

Références

Allen, L.F., Stimson, M.R., King, O.A., Norrad, R.E., Lucas, S.G., Mann, A., Hinds, S.J., Park, A.F., Calder, J.H., Maddin, H., and Montplaisir, M. 2022. A Batrachichnus salamandroides trackway from the Minto Formation of central New Brunswick, Canada: implications for alternative trace maker interpretations. Atlantic Geoscience, 58, pp. 239–260. https://doi.org/10.4138/atlgeol.2022.010 DOI: https://doi.org/10.4138/atlgeol.2022.010

Archer, A.W., Calder, J.H., Gibling, M.R., Naylor, R.D., Reid, D.R., and Wightman, W.G. 1995. Invertebrate trace fossils and agglutinated foraminifera as indicators of marine influence within the classic Carboniferous section at Joggins, Nova Scotia, Canada. Canadian Journal of Earth Sciences, 32 (12), pp. 2027–2039. https://doi.org/10.1139/e95-156 DOI: https://doi.org/10.1139/e95-156

Béthoux, O., Norrad, R.E., Stimson, M.R., King, O.A., Allen, L.F., Deregnaucourt, I., Hinds, S.J., Lewis, J.H., and Schneider, J.W. 2021. A unique, large-sized stem Odonata (Insecta) found in the early Pennsylvanian of New Brunswick (Canada). Fossil Record, 24 (2), pp. 207–221. https://doi.org/10.5194/fr-24-207-2021 DOI: https://doi.org/10.5194/fr-24-207-2021

Brandt, D.S. 2007. Multiple-Rusophycus (arthropod ichnofossil) assemblages and their significance. Ichnos, 15 (1), pp. 28–43. https://doi.org/10.1080/10420940701192898 DOI: https://doi.org/10.1080/10420940701192898

Briggs, D.E. 1979. A giant myriapod trail from the Namurian of Arran, Scotland. Palaeontology, 22 (2), pp. 273–291.

Briggs, D.G., Plint, A.G., and Pickerill, R.K. 1984. Arthropleura trails from the Westphalian of eastern Canada. Palaeontology, 27 (4), pp. 843–855.

Bromley, R. and Asgaard, U. 1979. Triassic freshwater ichnocoenoses from Carlsberg Fjord, East Greenland. Palaeogeography, Palaeoclimatology, Palaeoecology, 28, pp.39–80. https://doi.org/10.1016/0031-0182(79)90112-3 DOI: https://doi.org/10.1016/0031-0182(79)90112-3

Brooks, H.K. 1962. The Paleozoic Eumalacostraca of North America. Bulletins of American Paleontology, 44, pp. 163–335.

Buatois, L.A. and Mángano, M.G. 2003. Early colonization of the deep sea: ichnologic evidence of deep-marine benthic ecology from the Early Cambrian of northwest Argentina. Palaios, 18 (6), pp. 572–581. https://doi.org/10.1669/0883-1351(2003)018<0572:ECOTDS>2.0.CO;2 DOI: https://doi.org/10.1669/0883-1351(2003)018<0572:ECOTDS>2.0.CO;2

Buhler, P.B. and Grey, M. 2017. Xiphosuran digging traces at the late Carboniferous Joggins Fossil Cliffs UNESCO World Heritage Site, Nova Scotia, Canada. Ichnos, 24 (3), pp. 179–190. https://doi.org/10.1080/10420940.2016.1244055 DOI: https://doi.org/10.1080/10420940.2016.1244055

Calder, J.H. 1998. The Carboniferous evolution of Nova Scotia. Geological Society, London, Special Publications, 143 (1), pp. 261–302. https://doi.org/10.1144/GSL.SP.1998.143.01.19 DOI: https://doi.org/10.1144/GSL.SP.1998.143.01.19

Calder, J.H., Gibling, M.R., Scott, A.C., Davies, S.J., and Hebert, B.L. 2006. A fossil lycopsid forest succession in the classic Joggins section of Nova Scotia: paleoecology of a disturbance-prone Pennsylvanian wetland. Wetlands Through Time, 399, pp. 169–195. https://doi.org/10.1130/2006.2399(09) DOI: https://doi.org/10.1130/2006.2399(09)

Carpenter, D.K., Falcon‐Lang, H.J., Benton, M.J., and Grey, M. 2015. Early Pennsylvanian (Langsettian) fish assemblages from the Joggins Formation, Canada, and their implications for palaeoecology and palaeogeography. Palaeontology, 58 (4), pp. 661–690. https://doi.org/10.1111/pala.12164 DOI: https://doi.org/10.1111/pala.12164

Carroll, R.L. 1964. The earliest reptiles. Journal of the Linnean Society of London (Zoology), 45, pp. 61–83. https://doi.org/10.1111/j.1096-3642.1964.tb00488.x DOI: https://doi.org/10.1111/j.1096-3642.1964.tb00488.x

Carroll, R.L. 1966. Microsaurs from the Westphalian b of Joggins, Nova Scotia, Proceedings of the Linnean Society of London, 177 (1), pp. 63–97. https://doi.org/10.1111/j.1095-8312.1966.tb00952.x DOI: https://doi.org/10.1111/j.1095-8312.1966.tb00952.x

Carroll, R.L. 1967. Labyrinthodonts from the Joggins Formation. Journal of Paleontology, pp. 111–142.

Carroll, R.L., 2009. The rise of amphibians: 365 million years of evolution. Johns Hopkins University Press, 392 pp.

Crimes, T.P. and Herdman, J. 1970 Trilobite tracks and other trace fossils from the Upper Cambrian of North Wales. Geological Journal, 7 (1), pp. 47–68. https://doi.org/10.1002/gj.3350070104 DOI: https://doi.org/10.1002/gj.3350070104

Dafoe, L., Stimson, M.R., and Gibling, M.R. 2011. One more piece of the puzzle: new developments in the trace fossil record from the Pennsylvanian Joggins Formation. Atlantic Geoscience Society, Annual Colloquium 2011 Abstracts; also published in Atlantic Geology, 47, p. 17. https://id.erudit.org/iderudit/ageo47abs01

Davies, S.J., Gibling, M.R., Rygel, M.C., Calder, J.H., and Skilliter, D.M. 2005. The Pennsylvanian Joggins Formation of Nova Scotia: sedimentological log and stratigraphic framework of the historic fossil cliffs. Atlantic Geology, 41, pp. 115–142. https://doi.org/10.4138/182 DOI: https://doi.org/10.4138/182

Dawson, J.W. 1862. Notice of the discovery of additional remains of land animals in the Coal-Measures of the South Joggins, Nova Scotia. Quarterly Journal of the Geological Society of London, 18, pp. 5–8. https://doi.org/10.1144/GSL.JGS.1862.018.01-02.10 DOI: https://doi.org/10.1144/GSL.JGS.1862.018.01-02.10

Dawson, J.W. 1863. Air-breathers of the Coal Period: descriptive account of the remains of land animals found in the Coal Formation of Nova Scotia. Dawson Brothers, Montreal, 102 pp. https://doi.org/10.5962/bhl.title.167425 DOI: https://doi.org/10.5962/bhl.title.167425

Dawson, J.W. 1864. On the fossils of the genus Rusophycus. Canadian Naturalist and Geologist, 1, pp. 363–367.

Dawson, J.W. 1873. Note on footprints from the Carboniferous of Nova Scotia, in the collection of the Geological Survey of Canada. Geological Magazine, Series 1, 9, pp. 251–253. https://doi.org/10.1017/S0016756800465052 DOI: https://doi.org/10.1017/S0016756800465052

Dawson, J.W. 1891. On the mode of occurrence of remains of land animals in erect trees at South Joggins, Nova Scotia. Transactions of the Royal Society of Canada, Section IV, 9, pp. 127–128.

Dawson, J.W. 1894. Synopsis of the air-breathing animals of the Paleozoic in Canada, up to 1894. Transactions of the Royal Society of Canada, 12, Section IV, pp. 71–88.

Dernov, V.S. 2023. Hankoichnus ichnogen. nov., a new arthropod (?) trace fossil from the Car-boniferous of the Donets Basin (Ukraine). Geologìcnij žurnal/Geologiceskij žur-nal/Heolohichnyi Zhurnal, 1, pp. 53–58. https://doi.org/10.30836/igs.1025-6814.2023.1.265486 DOI: https://doi.org/10.30836/igs.1025-6814.2023.1.265486

Dietrich, J., Lavoie, D., Hannigan, P., Pinet, N., Castonguay, S., Giles, P., and Hamblin, A. 2011. Geological setting and resource potential of conventional petroleum plays in Paleozoic basins in eastern Canada. Bulletin of Canadian Petroleum Geology, 59 (1), pp. 54–84. https://doi.org/10.2113/gscpgbull.59.1.54 DOI: https://doi.org/10.2113/gscpgbull.59.1.54

Falcon-Lang, H.J. 2004. Pennsylvanian tropical rainforests responded to glacial–interglacial rhythms. Geology, 32, pp. 689-692. https://doi.org/10.1130/G20523.1 DOI: https://doi.org/10.1130/G20523.1

Falcon-Lang, H.J. 2006. A history of research at the Joggins Fossil Cliffs of Nova Scotia, Canada, the world's finest Pennsylvanian section. Proceedings of the Geologists' Association, 117 (4), pp. 377–392. https://doi.org/10.1016/S0016-7878(06)80044-1 DOI: https://doi.org/10.1016/S0016-7878(06)80044-1

Falcon-Lang, H.J., Benton, M.J., Braddy, S.J., and Davies, S.J. 2006. The Pennsylvanian tropical biome reconstructed from the Joggins Formation of Nova Scotia, Canada. Journal of the Geological Society, 163 (3), pp. 561–576. https://doi.org/10.1144/0016-764905-063 DOI: https://doi.org/10.1144/0016-764905-063

Falkingham, P.L., Bates, K.T., Avanzini, M., Bennett, M., Bordy, E.M., Breithaupt, B.H., Castanera, D., Citton, P., Díaz‐Martínez, I., Farlow, J.O., and Fiorillo, A.R. 2018. A standard protocol for documenting modern and fossil ichnological data. Palaeontology, 61 (4), pp. 469–480. https://doi.org/10.1111/pala.12373 DOI: https://doi.org/10.1111/pala.12373

Feldmann, R.M., Osgood, R.G.Jr., Szmuc, E.J. and Meinke, D.W. 1978. Chagrinichnites brooksi, a new trace fossil of arthropod origin. Journal of Paleontology, pp. 287–294.

Ferguson, L. 1966. The recovery of some large track-bearing slabs from Joggins, Nova Scotia. Maritime Sediments, 2, pp. 128–130. https://doi.org/10.4138/1501 DOI: https://doi.org/10.4138/1501

Gand, G., Garric, J., Schneider, J., Walter, H., Lapeyrie, J., Martin, C., and Thiery, A. 2008. Notostraca trackways in Permian playa environments of the Lodève basin (France). Journal of Iberian Geology, 34 (1), pp. 73–108.

Gibb, S. and Pemberton, S.G. 2017. Rusophycus by any other name is still Rusophycus carleyi. Ichnos, 24 (1), pp. 19–26. https://doi.org/10.1080/10420940.2016.1257486 DOI: https://doi.org/10.1080/10420940.2016.1257486

Gibling, M.R., Culshaw, N., Pascucci, V., Waldron, J.W.F., and Rygel, M.C. 2019. The Maritimes Basin of Atlantic Canada: basin creation and destruction during the Paleozoic assembly of Pangea. In The sedimentary basins of the United States and Canada. Elsevier, pp. 267–314. https://doi.org/10.1016/B978-0-444-63895-3.00006-1 DOI: https://doi.org/10.1016/B978-0-444-63895-3.00006-1

Grabau, A.W. and Shimer, H.W. 1910. North American index fossils. 2, pp. 818–819.

Gradstein, F.M. and Ogg, J.G. 2020. The chronostratigraphic scale. In Geologic time scale 2020. Elsevier, pp 21–32. https://doi.org/10.1016/B978-0-12-824360-2.00002-4 DOI: https://doi.org/10.1016/B978-0-12-824360-2.00002-4

Grey, M.G. and Finkel, Z.V. 2011. The Joggins Fossil Cliffs UNESCO World Heritage site: a review of recent research. Atlantic Geology, 47, pp. 185–200. https://doi.org/10.4138/atlgeol.2011.009 DOI: https://doi.org/10.4138/atlgeol.2011.009

Grey, M., Pufahl, P.K., and Aziz, A.A. 2011. Using multiple environmental proxies to determine degree of marine influence and paleogeographical position of the Joggins Fossil Cliffs, UNESCO World Heritage Site. Palaios, 26 (5), pp. 256–263. https://doi.org/10.2110/palo.2010.p10-118r DOI: https://doi.org/10.2110/palo.2010.p10-118r

Grey, M., Finkel, Z.V., Pufahl, P.K., and Reid, L.M. 2012. Evolutionary mode of the ostracod, Velatomorpha altilis, from the Joggins Fossil Cliffs UNESCO World Heritage Site. Lethaia, 45 (4), pp. 615–623. https://doi.org/10.1111/j.1502-3931.2012.00328.x DOI: https://doi.org/10.1111/j.1502-3931.2012.00328.x

Hall, J. 1852. Palaeontology of New-York: Containing descriptions of the organic remains of the lower middle division of the New-York system., Volume 2. C. Van Benthuysen, 362 pp.

Hardy, P.G. 1970. New xiphosurid trails from the Upper Carboniferous of northern England. Palaeontology, 13, pp. 188–190.

Heidtke, U. 1990. Pollichianum cubichnum nov. ichnogen. et ichnospec. und Pollichianum repichnum nov. ichnospec., zwei neue Invertebratenichnia aus dem Saar-Pfälzischen Rotliegend (Unter-Perm, SW-Deutschland). Mitteilungen der Pollichia, 77, pp. 133–139.

Hofmann, R., Mángano, M.G., Elicki, O., and Shinaq, R. 2012. Paleoecologic and biostratigraphic significance of trace fossils from shallow-to marginal-marine environments from the Middle Cambrian (Stage 5) of Jordan. Journal of Paleontology, 86 (6), pp. 931–955. https://doi.org/10.1666/11-129R1.1 DOI: https://doi.org/10.1666/11-129R1.1

Holmes, R.B., Carroll, R.L., and Reisz, R.R. 1998. The first articulated skeleton of Dendrerpeton acadianum (Temnospondyli, Dendrerpetonidae) from the Lower Pennsylvanian locality of Joggins, Nova Scotia, and a review of its relationships. Journal of Vertebrate Palaeontology, 18, pp. 64–79. https://doi.org/10.1080/02724634.1998.10011034 DOI: https://doi.org/10.1080/02724634.1998.10011034

Keighley, D.G. and Pickerill, R.K. 1996. Small Cruziana, Rusophycus, and related ichnotaxa from eastern Canada: the nomenclatural debate and systematic ichnology. Ichnos, 4 (4), pp. 261–285. https://doi.org/10.1080/10420949609380136 DOI: https://doi.org/10.1080/10420949609380136

King, O. A., Stimson, M.R., and Lucas, S.G. 2019. The ichnogenus Kouphichnium and related xiphosuran traces from the Steven C. Minkin Paleozoic Footprint Site (Union Chapel Mine), Alabama, USA: ichnotaxonomic and paleoenvironmental implications. Ichnos, 26 (4), pp. 266–302. https://doi.org/10.1080/10420940.2018.1561447 DOI: https://doi.org/10.1080/10420940.2018.1561447

Knecht, R.J., Benner, J.S., Rogers, D.C., and Ridge, J.C. 2009. Surculichnus bifurcauda n. igen., n. isp., a trace fossil from Late Pleistocene glaciolacustrine varves of the Connecticut River Valley, USA, attributed to notostracan crustaceans based on neoichnological experimentation. Palaeogeography, Palaeoclimatology, Palaeoecology, 272 (3-4), pp. 232–239. https://doi.org/10.1016/j.palaeo.2008.10.013 DOI: https://doi.org/10.1016/j.palaeo.2008.10.013

Lerner, A.J. and Lucas, S.G. 2015. A Selenichnites ichnoassociation from early Permian tidal flats of the prehistoric trackways national monument of south-central New Mexico. Bulletin of the New Mexico Museum of Natural History and Science, 65, pp. 141–152.

Letourmy, Y., Driese, S.G., and Sims, J.R. 2021. Absence of evidence of climate-driven cycles in Carboniferous deposits of Joggins, Nova Scotia, Canada: influence of salt withdrawal tectonics on deposition and pedogenesis. Journal of Sedimentary Research, 91 (2), pp. 167–185. https://doi.org/10.2110/jsr.2020.78 DOI: https://doi.org/10.2110/jsr.2020.78

Lyell, C. and Dawson, J.W. 1853. On the remains of a reptile (Dendrerpeton acadianum, Wyman and Owen) and of a land shell discovered in the interior of an erect fossil tree in the coal measures of Nova Scotia. Quarterly journal of the Geological Society, 9 (1–2), pp. 58–67. https://doi.org/10.1144/GSL.JGS.1853.009.01-02.20 DOI: https://doi.org/10.1144/GSL.JGS.1853.009.01-02.20

Mann, A., Gee, B.M., Pardo, J.D., Marjanović, D., Adams, G.R., Calthorpe, A.S., Maddin, H.C., and Anderson, J.S. 2020. Reassessment of historic ‘microsaurs’ from Joggins, Nova Scotia, reveals hidden diversity in the earliest amniote ecosystem. Papers in Palaeontology, 6 (4), pp. 605–625. https://doi.org/10.1002/spp2.1316 DOI: https://doi.org/10.1002/spp2.1316

Marchetti, L., Belvedere, M., Voigt, S., Klein, H., Castanera, D., Díaz-Martínez, I., Marty, D., Xing, L., Feola, S., Melchor, R.N., and Farlow, J.O. 2019. Defining the morphological quality of fossil footprints. Problems and principles of preservation in tetrapod ichnology with examples from the Palaeozoic to the present. Earth Science Reviews, 193, pp. 109–145. https://doi.org/10.1016/j.earscirev.2019.04.008 DOI: https://doi.org/10.1016/j.earscirev.2019.04.008

Marchetti, L., Voigt, S., Lucas, S.G., Stimson, M.R., King, O.A., and Calder, J.H. 2020 Footprints of the earliest reptiles. Notalacerta missouriensis ― ichnotaxonomy; potential trackmakers, biostratigrapy, palaeobiogeography, and palaeoecology. Annales Societatis Geologorum Poloniae, 90, pp. 271–29. https://doi.org/10.14241/asgp.2020.13 DOI: https://doi.org/10.14241/asgp.2020.13

Marchetti, L., Voigt, S., Buchwitz, M., MacDougall, M.J., Lucas, S.G., Fillmore, D.L., Stimson, M.R., King, O.A., Calder, J.H., and Fröbisch, J. 2021. Tracking the origin and early evolution of reptiles. Frontiers in Ecology and Evolution, 9, pp. 385. https://doi.org/10.3389/fevo.2021.696511 DOI: https://doi.org/10.3389/fevo.2021.696511

Matthew, G.F. 1903. On batrachian and other footprints from the Coal Measures of Joggins, N.S. Natural History Society of New Brunswick Bulletin, 21, pp. 103–108.

Meade, L., Jones, A., and Butler R. 2016. A revision of tetrapod footprints from the late Carboniferous of the West Midlands, UK. Peerj, e2718, pp. 1–18. https://doi.org/10.7717/peerj.2718 DOI: https://doi.org/10.7717/peerj.2718

Melchor, R.N. and Cardonatto, M.C. 2014. Insights on the behavior of late Paleozoic aquatic crustaceans (Pygocephalomorpha?): compound trace fossils from western Argentina. Ichnos, 21 (2), pp. 76-99. https://doi.org/10.1080/10420940.2013.879868 DOI: https://doi.org/10.1080/10420940.2013.879868

Milàn, J., Falkingham, P.L., Marty, D., and Richter, A. 2016. Chapter 1. Experimental and comparative ichnology. In Dinosaur tracks: the next steps. Edited by P.L. Falkinham, D. Marty, and A. Richter. Indiana University Press, pp. 14–27.

Miller, M.F. 1982. Limulicubichnus: a new ichnogenus of limulid resting traces. Journal of Paleontology, pp. 429–433.

Minter, N.J. and Braddy, S.J. 2009. Ichnology of an early Permian intertidal flat: the Robledo Mountains Formation of southern New Mexico, USA. Special Papers in Palaeontology, 82 (82), pp. 5–107.

O’Brien, L.J., Braddy, S.J., and Radley, J.D. 2009. A new arthropod resting trace and associated suite of trace fossils from the Lower Jurassic of Warwickshire, England. Palaeontology, 52 (5), pp. 1099–1112. https://doi.org/10.1111/j.1475-4983.2009.00901.x DOI: https://doi.org/10.1111/j.1475-4983.2009.00901.x

Petti, F.M., Avanzini, M., Belvedere, M., De Gasperi, M., Ferretti, P., Girardi, S., Remondino, F., and Tomasoni, R. 2008. Digital 3-D modelling of dinosaur footprints by photogrammetry and laser scanning techniques: integrated approach at the Coste dell’Anglone tracksite (Lower Jurassic, southern Alps, northern Italy). Acta Geologicav, 83, pp. 303–315.

Pickerill, R.K. 1992. Carboniferous nonmarine invertebrate ichnocoenoses from southern New Brunswick, eastern Canada. Ichnos, 2 (1), pp. 21–35. https://doi.org/10.1080/10420949209380072 DOI: https://doi.org/10.1080/10420949209380072

Prescott, Z.M., Stimson, M.R., Dafoe, L.T., Gibling, M.R., MacRae, R.A., Calder, J.H., and Hebert, B.L. 2014. Microbial mats and ichnofauna of a fluvial-tidal channel in the Lower Pennsylvanian Joggins Formation, Canada. Palaios, 29 (12), pp. 624–645. https://doi.org/10.2110/palo.2013.073 DOI: https://doi.org/10.2110/palo.2013.073

Rmich, A., Lagnaoui, A., Hminna, A., Saber, H., Zouheir, T., and Lallensack, J.N. 2023. Captorhinid trackways from mid to late Permian red beds in Morocco: implications for locomotion and the palaeobiogeography of northwest Gondwana. Palaeogeography, Palaeoclimatology, Palaeoecology, 111700, pp. 1-21. https://doi.org/10.1016/j.palaeo.2023.111700 DOI: https://doi.org/10.1016/j.palaeo.2023.111700

Romano, M. and Whyte, M.A. 1987. A limulid trace fossil from the Scarborough Formation (Jurassic) of Yorkshire; its occurrence, taxonomy and interpretation. Proceedings of the Yorkshire Geological Society, 46 (2), pp. 85–95. https://doi.org/10.1144/pygs.46.2.85 DOI: https://doi.org/10.1144/pygs.46.2.85

Romano, M. and Whyte, M. 1990. Selenichnites, a new name for the ichnogenus Selenichnus Romano & Whyte, 1987. Proceedings of the Yorkshire Geological Society, 48 (2), pp. 221–221. https://doi.org/10.1144/pygs.48.2.221 DOI: https://doi.org/10.1144/pygs.48.2.221

Romano, M. and Whyte, M.A. 2015. A review of the trace fossil Selenichnites. Proceedings of the Yorkshire Geological Society, 60 (4), pp. 275–288. https://doi.org/10.1144/pygs2015-357 DOI: https://doi.org/10.1144/pygs2015-357

Romilio, A. and Salisbury, S.W. 2014. Large dinosaurian tracks from the upper Cretaceous (Cenomanian–Turonian) portion of the Winton Formation, Lark Quarry, central-western Queensland, Australia: 3D photogrammetric analysis renders the “stampede trigger’ scenario unlikely. Cretaceous Research. v. 51, pp. 186–207. https://doi.org/10.1016/j.cretres.2014.06.003 DOI: https://doi.org/10.1016/j.cretres.2014.06.003

Ryan, R.J. 1986. Fossil myriapod trails in the Permo-Carboniferous strata of northern Nova Scotia, Canada. Atlantic Geology, 22 (2), pp. 156–161. https://doi.org/10.4138/1604 DOI: https://doi.org/10.4138/1604

Rygel M.C. and Gibling M.R. 2006, Natural geomorphic variability recorded in a high-accommodation setting: fluvial architecture of the Pennsylvanian Joggins Formation of Atlantic Canada: Journal of Sedimentary Research, 76, pp. 1230–1251. https://doi.org/10.2110/jsr.2006.100 DOI: https://doi.org/10.2110/jsr.2006.100

Rygel, M.C., Sheldon, E.P., Stimson, M.R., Calder, J.H., Ashley, K.T., and Salg, J.L. 2014. The Pennsylvanian Springhill Mines Formation: sedimentological framework for a portion of the Joggins Fossil Cliffs UNESCO World Heritage Site. Atlantic Geology, 50, pp. 249–289. https://doi.org/10.4138/atlgeol.2014.013 DOI: https://doi.org/10.4138/atlgeol.2014.013

Savage, N.M. 1971. A varvite ichnocoenosis from the Dwyka Series of Natal. Lethaia, 4 (2), pp. 217–233. https://doi.org/10.1111/j.1502-3931.1971.tb01290.x DOI: https://doi.org/10.1111/j.1502-3931.1971.tb01290.x

Schneider, J.W., Lucas, S.G., Werneburg, R., Rößler, R., and Spielmann, J.A. 2010. Euramerican Late Pennsylvanian/Early Permian arthropleurid/tetrapod associations–implications for the habitat and paleobiology of the largest terrestrial arthropod. In Carboniferous–Permian transition in Canon del Cobre, northern New Mexico. Edited by S.G. Lucas, J.W. Schneider, and J. Spielmann. New Mexico Museum Nature History and Science Bulletin, 49, pp. 49–70.

Steen, M.C. 1934. The amphibian fauna from the South Joggins, Nova Scotia. In Proceedings of the Zoological Society of London, pp. 465–504. https://doi.org/10.1111/j.1096-3642.1934.tb01644.x DOI: https://doi.org/10.1111/j.1096-3642.1934.tb01644.x

Stimson, M. and MacRae, R.A. J.H. 2010. Evidence of fossil horseshoe crabs from Joggins, Nova Scotia: paleoichnology and paleoenvironmental implications. Atlantic Geology, 46 (1), p. 69. https://id.erudit.org/iderudit/ageo46abs01

Stimson, M., Lucas, S.G., and Melanson, G. 2012. The smallest known tetrapod footprints: Batrachichnus salamandroides from the Carboniferous of Joggins, Nova Scotia, Canada. Ichnos, 19 (3), pp. 127–140. https://doi.org/10.1080/10420940.2012.685206 DOI: https://doi.org/10.1080/10420940.2012.685206

Stimson, M.R., Miller, R.F., and Lucas, S.G. 2016. Reassignment of vertebrate ichnotaxa from the Upper Carboniferous ‘Fern Ledges’, Lancaster Formation, Saint John, New Brunswick, Canada. Atlantic Geology, 52, pp. 20–34. https://doi.org/10.4138/atlgeol.2016.002 DOI: https://doi.org/10.4138/atlgeol.2016.002

Teichert, C. 1948. A simple device for coating fossils with ammonium chloride. Journal of Paleontology, 22, pp. 102–104.

Trewin, N.H. 1994. A draft system for the identification and description of arthropod trackways. Palaeontology, 37 (4), pp. 811-823.

Ulrich, E. O. and Bassler, R. S. 1923. Paleozoic Ostracoda: their morphology, classification and occurrence. Maryland Geological Survey, Silurian, 9. pp. 271–391.

Utting, J., Giles, P.S., and Dolby, G. 2010. Palynostratigraphy of Mississippian and Pennsylvanian rocks, Joggins area, Nova Scotia and New Brunswick, Canada. Palynology, 34 (1), pp. 43–89. https://doi.org/10.1080/01916121003620569 DOI: https://doi.org/10.1080/01916121003620569

Vallon, L.H. and Roeper, S. 2006. Tripartichnus n. igen. ― a new trace fossil from the Buntsandstein (Lower Triassic) and from the Solnhofen lithographic limestones (Upper Jurassic), Germany: Tripartichnus n. igen. ― ein neues Spurenfossil aus dem Buntsandstein (Untertrias) und dem Solnhofener Plattenkalk (Oberjura), Deutschland. Paläontologische Zeitschrift, 80 (2), p. 156. https://doi.org/10.1007/BF02988974 DOI: https://doi.org/10.1007/BF02988974

Waldron, J.W. and Rygel, M.C. 2005. Role of evaporite withdrawal in the preservation of a unique coal-bearing succession: Pennsylvanian Joggins Formation, Nova Scotia. Geology, 33 (5), pp. 337–340. https://doi.org/10.1130/G21302.1 DOI: https://doi.org/10.1130/G21302.1

Waldron, J.W., Rygel, M.C., Gibling, M.R., and Calder, J.H. 2013. Evaporite tectonics and the late Paleozoic stratigraphic development of the Cumberland basin, Appalachians of Atlantic Canada. Geological Society of America Bulletin, 125 (5–6), pp. 945–960. https://doi.org/10.1130/B30718.1 DOI: https://doi.org/10.1130/B30718.1

Williams, E.P. 1974. Geology and petroleum possibilities in and around Gulf of St. Lawrence. American Association of Petroleum Geology Bulletin, 58 (6), pp. 1137–1155. https://doi.org/10.1306/83D9162D-16C7-11D7-8645000102C1865D DOI: https://doi.org/10.1306/83D9162D-16C7-11D7-8645000102C1865D

Zatoń, M., Grey, M., and Vinn, O. 2014. Microconchid tubeworms (class Tentaculita) from the Joggins Formation (Pennsylvanian), Nova Scotia, Canada. Canadian Journal of Earth Sciences, 51 (7), pp. 669–676. https://doi.org/10.1139/cjes-2014-0061 DOI: https://doi.org/10.1139/cjes-2014-0061

Publié-e

2024-08-30

Comment citer

Malay, B. C., Stimson, M. R., King, O. A., Faulkner, S., Hebert, B. L., Calder, J. H., … Lucas, S. G. (2024). Nouvel ichnogenre et nouvelle ichnoespèce de cubichnium, le Pygocephalichnium reidi, du site du patrimoine mondial de l’UNESCO des falaises fossilifères de Joggins, au Canada, et ichnotaxons connexes. Atlantic Geoscience, 60, 185–203. https://doi.org/10.4138/atlgeo.2024.009

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